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The notion that while there isn't necessarily something wrong with someone/something, the two elements aren't a match.
"Dude, why did y'all break up?"
"Classic Mona Lisa Principle. She was beautiful, mysterious, and intriguing, but (pointing to my brain) didn't match a damn thing in my house."
When a cleavage looks kind of perky and happy but also a little on the sad any unperky side at the same time.
"Have you seen Emily's breasts? Her tits are like "Wow!" but also kind of "Meh." at the same time... I feel like her cleavage is somehow smiling at me--but also frowning at me! It's like she's got that whole Mona Lisa Cleavage thing going on. You know, like auntie Sara."