Balls; courage; brutish masculinity. Fr. Italian "coglioni". Cf. Sp. cojones, Fr. cuillons; see break my balls.
You need a giant set of culliones to talk to Tony like that.
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Sussex 1950's for testes, may derive from cojones, Sanish, same meaning.
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wen running or dancing or just walking yur balls swing and hit yur thighs and or ass(if big balls like james eagle's)
james had captian cullions while running in cricket